The circulatory system
consists of longitudinal and transverse blood vessels, in which blood flows
continuously. The circulatory system is complicated and the arrangement of
blood vessels behind the 14th segment is different from the arrangement from
the first 14th segments. So a general plan of circulatory system in earthworm
is being discussed.
There are three main
longitudinal blood vessels, besides smaller vessels, 4/5 pairs of hearts found
in earthworm:
Dorsal Vessel
It runs throughout the
length of the body in the mid dorsal line. It is present above the alimentary
canal. Its wall is thick and muscular. It shows rhythmical peristaltic
contraction from behind to the front. The backward flow is prevented by a pair
of valve present in front of each septum (septum is partition wall between the
segments). It serves only as a collecting vessel. It receives blood from
intestine, nephridia; body wall etc. in the first 14th segment, the dorsal
vessel becomes a distributing vessel. It supplies blood to anterior region, of
the earthworm. It also sends blood to the four pairs of hearts. The hearts pump
the blood into the ventral vessel.
Ventral Vessel
It runs throughout the
length of the body below the alimentary canal. It is a distributing vessel
only. The blood flows from anterior to posterior end. Valves are absent in its
lumen (space). A number of small blood vessels are given off, which distribute
blood to the body wall, septa and nephridia of each segment.
Sub-Neural Vessel
It extends along the
mid-ventral line of the body wall below the nerve cord and extends from 14th
segment backwards. In each segment, it collects blood from the nerve cord and
from ventral region of the body wall and the flow of blood is backward. Sub
neural vessel in the 14th segment divides into two lateral oesophageal vessels.
Sub-neural vessel communicates with the dorsal blood vessel through commissural
vessel.
Supra Oesophageal Vessel
It runs from 9th to
13th segment, between dorsal vessel and the oesophagus. At posterior it is
connected with the ventral vessel through the two latero-oesophagal hearts.
Hearts
The number of hearts
depends on type of the species. It maybe 4 or 5 pairs. Hearts are situated in
the 7th to 13th segments. They are also called aortic arch. The heart pumps
blood from the dorsal to the ventral vessel. The flow of blood cannot be reversed
due to the arrangement of the valves at the junction of dorsal vessel and
hearts.
Valves
The direction of flow
of blood is controlled by valves. They are present in the (1) Dorsal vessel (2)
Heart (3) at the points of junction of the chief vessels with the dorsal
vessels.
Blood
It is bright red in
color due to the presence of hemoglobin, which is dissolved in plasma. There
are also corpuscles in the blood which are colorless and nucleated.
Distribution of Food
The branches of the
ventral vessels that enter the wall of the digestive tract also break up into
innumerable capillaries. The product of digestion such as amino acids and
glucose enters the capillaries. The capillaries of the intestinal wall join to
the dorsal vessel. Thus the end products of digestion are distributed to all
parts of the body.
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